About Me
Dr. Fortune has been interested in evolutionary biology since childhood. He is passionate about understanding the mechanisms and evolution of animal behavior.
Education
Ph.D.
; University of Chicago
; Biology
; 1995
B.S. ; University of Chicago ; ; 1989
B.S. ; University of Chicago ; ; 1989
Website
2025 Fall Courses
BIOL 790E - DOCTORAL DISSERTATION
BIOL 726 - INDEPENDENT STUDY II
BIOL 790A - DOCT DISSERTATION & RESRCH
BIOL 790C - DOCTORAL DISSERTN & RESRCH
BIOL 700B - MASTER'S PROJECT
BIOL 725 - INDEPENDENT STUDY I
BIOL 790B - DOCT DISSERTATION & RESRCH
BIOL 790D - DOCT DISSERTATION & RESRCH
BIOL 491 - SENIOR PROJECT
BIOL 492 - RESEARCH & INDEPENDENT STUDY
BIOL 792B - PRE-DOCTORAL RESEARCH
BIOL 630 - CRITICAL THINKING FOR LIFE SCI
BIOL 792C - PRE-DOCTORAL RESEARCH
BIOL 726 - INDEPENDENT STUDY II
BIOL 790A - DOCT DISSERTATION & RESRCH
BIOL 790C - DOCTORAL DISSERTN & RESRCH
BIOL 700B - MASTER'S PROJECT
BIOL 725 - INDEPENDENT STUDY I
BIOL 790B - DOCT DISSERTATION & RESRCH
BIOL 790D - DOCT DISSERTATION & RESRCH
BIOL 491 - SENIOR PROJECT
BIOL 492 - RESEARCH & INDEPENDENT STUDY
BIOL 792B - PRE-DOCTORAL RESEARCH
BIOL 630 - CRITICAL THINKING FOR LIFE SCI
BIOL 792C - PRE-DOCTORAL RESEARCH
Teaching Interests
Animal behavior and neuroscience
Past Courses
BIOL 150: LIVING IN A VARIABLE UNIVERSE
BIOL 200: CONCEPTS IN BIOLOGY
BIOL 337: COLLECTIVE INTEL IN BIOL SYST
BIOL 340: MAMMALIAN PHYSIOLOGY
BIOL 383: NEURAL BASIS OF BEHAVIOR
BIOL 447: CELLULAR&SYSTEMS NEUROSCIENCE
BIOL 495: HONORS SEMINAR IN BIOLOGY
BIOL 612: COMPARATIVE ANIMAL PHYSIOLOGY
BIOL 630: CRITICAL THINKING FOR LIFE SCI
BIOL 630: CRITICAL THINKING FOR THE LIFE SCIENCES
BIOL 200: CONCEPTS IN BIOLOGY
BIOL 337: COLLECTIVE INTEL IN BIOL SYST
BIOL 340: MAMMALIAN PHYSIOLOGY
BIOL 383: NEURAL BASIS OF BEHAVIOR
BIOL 447: CELLULAR&SYSTEMS NEUROSCIENCE
BIOL 495: HONORS SEMINAR IN BIOLOGY
BIOL 612: COMPARATIVE ANIMAL PHYSIOLOGY
BIOL 630: CRITICAL THINKING FOR LIFE SCI
BIOL 630: CRITICAL THINKING FOR THE LIFE SCIENCES
Research Interests
Mechanisms of Animal Behavior
Journal Article
Horacio G. Rotstein, David Sant, Alfred Amendolara, Eric S. Fortune.
2023. "LSTM-Based Recurrent Neural Network Provides Effective Short Term Flu Forecasting."
BMC Publich Health , vol. 23 , pp. 1788.
I Uyanik, S Sefati, S A Stamper, K A Cho, M M Ankarali, Eric S. Fortune, N J Cowan. 2020. "Variability in locomotor dynamics reveals the critical role of feedback in task control.." eLife , vol. 9 .
I Uyanik, S A Stamper, N J Cowan, Eric S. Fortune. 2019. "Sensory Cues Modulate Smooth Pursuit and Active Sensing Movements.." Frontiers in behavioral neuroscience , vol. 13 , pp. 59.
J E Elie, S Hoffmann, J L Dunning, M J Coleman, Eric S. Fortune, J F Prather. 2019. "From Perception to Action: The Role of Auditory Input in Shaping Vocal Communication and Social Behaviors in Birds.." Brain, behavior and evolution , vol. 94 , no. 1-4 , pp. 51-60.
Melissa Coleman, Eric S. Fortune. 2018. "Quick Guide: Duet singing in plain-tailed wrens." Current Biology , vol. 28 , no. 11 , pp. 643-645.
I Uyanik, S Sefati, S A Stamper, K A Cho, M M Ankarali, Eric S. Fortune, N J Cowan. 2020. "Variability in locomotor dynamics reveals the critical role of feedback in task control.." eLife , vol. 9 .
I Uyanik, S A Stamper, N J Cowan, Eric S. Fortune. 2019. "Sensory Cues Modulate Smooth Pursuit and Active Sensing Movements.." Frontiers in behavioral neuroscience , vol. 13 , pp. 59.
J E Elie, S Hoffmann, J L Dunning, M J Coleman, Eric S. Fortune, J F Prather. 2019. "From Perception to Action: The Role of Auditory Input in Shaping Vocal Communication and Social Behaviors in Birds.." Brain, behavior and evolution , vol. 94 , no. 1-4 , pp. 51-60.
Melissa Coleman, Eric S. Fortune. 2018. "Quick Guide: Duet singing in plain-tailed wrens." Current Biology , vol. 28 , no. 11 , pp. 643-645.
SHOW MORE
D Biswas, L A Arend, S A Stamper, B P Vágvölgyi, Eric S. Fortune, N J Cowan.
2018. "Closed-Loop Control of Active Sensing Movements Regulates Sensory Slip.."
Current biology : CB , vol. 28 , no. 24 , pp. 4029-4036.e4.
M S Madhav, R P Jayakumar, A Demir, S A Stamper, Eric S. Fortune, N J Cowan. 2018. "High-resolution behavioral mapping of electric fishes in Amazonian habitats.." Scientific reports , vol. 8 , no. 1 , pp. 5830.
E E Sutton, A Demir, S A Stamper, Eric S. Fortune, N J Cowan. 2016. "Dynamic modulation of visual and electrosensory gains for locomotor control.." Journal of the Royal Society, Interface , vol. 13 , no. 118 .
M L Rouse, Jr, T J Stevenson, Eric S. Fortune, G F Ball. 2015. "Reproductive state modulates testosterone-induced singing in adult female European starlings (Sturnus vulgaris).." Hormones and behavior , vol. 72 , pp. 78-87.
N J Cowan, M M Ankarali, J P Dyhr, M S Madhav, E Roth, S Sefati, S Sponberg, S A Stamper, Eric S. Fortune, T L Daniel. 2014. "Feedback control as a framework for understanding tradeoffs in biology.." Integrative and comparative biology , vol. 54 , no. 2 , pp. 223-37.
S Sefati, I D Neveln, E Roth, T R Mitchell, J B Snyder, M A Maciver, Eric S. Fortune, N J Cowan. 2013. "Mutually opposing forces during locomotion can eliminate the tradeoff between maneuverability and stability.." Proceedings of the National Academy of Sciences of the United States of America , vol. 110 , no. 47 , pp. 18798-803.
M S Madhav, S A Stamper, Eric S. Fortune, N J Cowan. 2013. "Closed-loop stabilization of the Jamming Avoidance Response reveals its locally unstable and globally nonlinear dynamics.." The Journal of experimental biology , vol. 216 , no. Pt 22 , pp. 4272-84.
S A Stamper, Eric S. Fortune, M J Chacron. 2013. "Perception and coding of envelopes in weakly electric fishes.." The Journal of experimental biology , vol. 216 , no. Pt 13 , pp. 2393-402.
S A Stamper, M S Madhav, N J Cowan, Eric S. Fortune. 2012. "Beyond the Jamming Avoidance Response: weakly electric fish respond to the envelope of social electrosensory signals.." The Journal of experimental biology , vol. 215 , no. Pt 23 , pp. 4196-207.
S A Stamper, E Roth, N J Cowan, Eric S. Fortune. 2012. "Active sensing via movement shapes spatiotemporal patterns of sensory feedback.." The Journal of experimental biology , vol. 215 , no. Pt 9 , pp. 1567-74.
P McGillivray, K Vonderschen, Eric S. Fortune, M J Chacron. 2012. "Parallel coding of first- and second-order stimulus attributes by midbrain electrosensory neurons.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 32 , no. 16 , pp. 5510-24.
Eric S. Fortune, C Rodríguez, D Li , G F Ball, M J Coleman. 2011. "Neural mechanisms for the coordination of duet singing in wrens.." Science (New York, N.Y.) , vol. 334 , no. 6056 , pp. 666-70.
N Khosravi-Hashemi, Eric S. Fortune, M J Chacron. 2011. "Coding movement direction by burst firing in electrosensory neurons.." Journal of neurophysiology , vol. 106 , no. 4 , pp. 1954-68.
E Roth, K Zhuang, S A Stamper, Eric S. Fortune, N J Cowan. 2011. "Stimulus predictability mediates a switch in locomotor smooth pursuit performance for Eigenmannia virescens.." The Journal of experimental biology , vol. 214 , no. Pt 7 , pp. 1170-80.
M J Chacron, Eric S. Fortune. 2010. "Subthreshold membrane conductances enhance directional selectivity in vertebrate sensory neurons.." Journal of neurophysiology , vol. 104 , no. 1 , pp. 449-62.
S A Stamper, E Carrera-G, E W Tan, V Fugère, R Krahe, Eric S. Fortune. 2010. "Species differences in group size and electrosensory interference in weakly electric fishes: implications for electrosensory processing.." Behavioural brain research , vol. 207 , no. 2 , pp. 368-76.
M J Chacron, N Toporikova, Eric S. Fortune. 2009. "Differences in the time course of short-term depression across receptive fields are correlated with directional selectivity in electrosensory neurons.." Journal of neurophysiology , vol. 102 , no. 6 , pp. 3270-9.
Eric S. Fortune, M J Chacron. 2009. "From molecules to behavior: organismal-level regulation of ion channel trafficking.." PLoS biology , vol. 7 , no. 9 , pp. e1000211.
E M Hitschfeld, S A Stamper, K Vonderschen, Eric S. Fortune, M J Chacron. 2009. "Effects of restraint and immobilization on electrosensory behaviors of weakly electric fish.." ILAR journal , vol. 50 , no. 4 , pp. 361-72.
S Carver, E Roth, N J Cowan, Eric S. Fortune. 2008. "Synaptic plasticity can produce and enhance direction selectivity.." PLoS computational biology , vol. 4 , no. 2 , pp. e32.
N J Cowan, Eric S. Fortune. 2007. "The critical role of locomotion mechanics in decoding sensory systems.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 27 , no. 5 , pp. 1123-8.
J U Ramcharitar, E W Tan, Eric S. Fortune. 2006. "Global electrosensory oscillations enhance directional responses of midbrain neurons in eigenmannia.." Journal of neurophysiology , vol. 96 , no. 5 , pp. 2319-26.
Eric S. Fortune. 2006. "The decoding of electrosensory systems.." Current opinion in neurobiology , vol. 16 , no. 4 , pp. 474-80.
Eric S. Fortune, G J Rose, M Kawasaki. 2006. "Encoding and processing biologically relevant temporal information in electrosensory systems.." Journal of comparative physiology. A, Neuroethology, sensory, neural, and behavioral physiology , vol. 192 , no. 6 , pp. 625-35.
E W Tan, J M Nizar, E Carrera-G, Eric S. Fortune. 2005. "Electrosensory interference in naturally occurring aggregates of a species of weakly electric fish, Eigenmannia virescens.." Behavioural brain research , vol. 164 , no. 1 , pp. 83-92.
J U Ramcharitar, E W Tan, Eric S. Fortune. 2005. "Effects of global electrosensory signals on motion processing in the midbrain of Eigenmannia.." Journal of comparative physiology. A, Neuroethology, sensory, neural, and behavioral physiology , vol. 191 , no. 9 , pp. 865-72.
Eric S. Fortune, G J Rose. 2003. "Voltage-gated Na+ channels enhance the temporal filtering properties of electrosensory neurons in the torus.." Journal of neurophysiology , vol. 90 , no. 2 , pp. 924-9.
Eric S. Fortune, G J Rose. 2002. "Roles for short-term synaptic plasticity in behavior.." Journal of physiology, Paris , vol. 96 , no. 5-6 , pp. 539-45.
Eric S. Fortune, G J Rose. 2001. "Short-term synaptic plasticity as a temporal filter.." Trends in neurosciences , vol. 24 , no. 7 , pp. 381-5.
Eric S. Fortune, G J Rose. 2000. "Short-term synaptic plasticity contributes to the temporal filtering of electrosensory information.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 20 , no. 18 , pp. 7122-30.
G J Rose, Eric S. Fortune. 1999. "Frequency-dependent PSP depression contributes to low-pass temporal filtering in Eigenmannia.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 19 , no. 17 , pp. 7629-39.
G J Rose, Eric S. Fortune. 1999. "Mechanisms for generating temporal filters in the electrosensory system.." The Journal of experimental biology , vol. 202 , no. Pt 10 , pp. 1281-9.
Eric S. Fortune, G J Rose. 1997. "Passive and active membrane properties contribute to the temporal filtering properties of midbrain neurons in vivo.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 17 , no. 10 , pp. 3815-25.
Eric S. Fortune, G J Rose. 1997. "Temporal filtering properties of ampullary electrosensory neurons in the torus semicircularis of Eigenmannia: evolutionary and computational implications.." Brain, behavior and evolution , vol. 49 , no. 6 , pp. 312-23.
G J Rose, Eric S. Fortune. 1996. "New techniques for making whole-cell recordings from CNS neurons in vivo.." Neuroscience research , vol. 26 , no. 1 , pp. 89-94.
Eric S. Fortune, D Margoliash. 1995. "Parallel pathways and convergence onto HVc and adjacent neostriatum of adult zebra finches (Taeniopygia guttata).." The Journal of comparative neurology , vol. 360 , no. 3 , pp. 413-41.
D Margoliash, Eric S. Fortune, M L Sutter, A C Yu, B D Wren-Hardin, A Dave. 1994. "Distributed representation in the song system of oscines: evolutionary implications and functional consequences.." Brain, behavior and evolution , vol. 44 , no. 4-5 , pp. 247-64.
Eric S. Fortune, D Margoliash. 1992. "Cytoarchitectonic organization and morphology of cells of the field L complex in male zebra finches (Taenopygia guttata).." The Journal of comparative neurology , vol. 325 , no. 3 , pp. 388-404.
D Margoliash, Eric S. Fortune. 1992. "Temporal and harmonic combination-sensitive neurons in the zebra finch's HVc.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 12 , no. 11 , pp. 4309-26.
M S Madhav, R P Jayakumar, A Demir, S A Stamper, Eric S. Fortune, N J Cowan. 2018. "High-resolution behavioral mapping of electric fishes in Amazonian habitats.." Scientific reports , vol. 8 , no. 1 , pp. 5830.
E E Sutton, A Demir, S A Stamper, Eric S. Fortune, N J Cowan. 2016. "Dynamic modulation of visual and electrosensory gains for locomotor control.." Journal of the Royal Society, Interface , vol. 13 , no. 118 .
M L Rouse, Jr, T J Stevenson, Eric S. Fortune, G F Ball. 2015. "Reproductive state modulates testosterone-induced singing in adult female European starlings (Sturnus vulgaris).." Hormones and behavior , vol. 72 , pp. 78-87.
N J Cowan, M M Ankarali, J P Dyhr, M S Madhav, E Roth, S Sefati, S Sponberg, S A Stamper, Eric S. Fortune, T L Daniel. 2014. "Feedback control as a framework for understanding tradeoffs in biology.." Integrative and comparative biology , vol. 54 , no. 2 , pp. 223-37.
S Sefati, I D Neveln, E Roth, T R Mitchell, J B Snyder, M A Maciver, Eric S. Fortune, N J Cowan. 2013. "Mutually opposing forces during locomotion can eliminate the tradeoff between maneuverability and stability.." Proceedings of the National Academy of Sciences of the United States of America , vol. 110 , no. 47 , pp. 18798-803.
M S Madhav, S A Stamper, Eric S. Fortune, N J Cowan. 2013. "Closed-loop stabilization of the Jamming Avoidance Response reveals its locally unstable and globally nonlinear dynamics.." The Journal of experimental biology , vol. 216 , no. Pt 22 , pp. 4272-84.
S A Stamper, Eric S. Fortune, M J Chacron. 2013. "Perception and coding of envelopes in weakly electric fishes.." The Journal of experimental biology , vol. 216 , no. Pt 13 , pp. 2393-402.
S A Stamper, M S Madhav, N J Cowan, Eric S. Fortune. 2012. "Beyond the Jamming Avoidance Response: weakly electric fish respond to the envelope of social electrosensory signals.." The Journal of experimental biology , vol. 215 , no. Pt 23 , pp. 4196-207.
S A Stamper, E Roth, N J Cowan, Eric S. Fortune. 2012. "Active sensing via movement shapes spatiotemporal patterns of sensory feedback.." The Journal of experimental biology , vol. 215 , no. Pt 9 , pp. 1567-74.
P McGillivray, K Vonderschen, Eric S. Fortune, M J Chacron. 2012. "Parallel coding of first- and second-order stimulus attributes by midbrain electrosensory neurons.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 32 , no. 16 , pp. 5510-24.
Eric S. Fortune, C Rodríguez, D Li , G F Ball, M J Coleman. 2011. "Neural mechanisms for the coordination of duet singing in wrens.." Science (New York, N.Y.) , vol. 334 , no. 6056 , pp. 666-70.
N Khosravi-Hashemi, Eric S. Fortune, M J Chacron. 2011. "Coding movement direction by burst firing in electrosensory neurons.." Journal of neurophysiology , vol. 106 , no. 4 , pp. 1954-68.
E Roth, K Zhuang, S A Stamper, Eric S. Fortune, N J Cowan. 2011. "Stimulus predictability mediates a switch in locomotor smooth pursuit performance for Eigenmannia virescens.." The Journal of experimental biology , vol. 214 , no. Pt 7 , pp. 1170-80.
M J Chacron, Eric S. Fortune. 2010. "Subthreshold membrane conductances enhance directional selectivity in vertebrate sensory neurons.." Journal of neurophysiology , vol. 104 , no. 1 , pp. 449-62.
S A Stamper, E Carrera-G, E W Tan, V Fugère, R Krahe, Eric S. Fortune. 2010. "Species differences in group size and electrosensory interference in weakly electric fishes: implications for electrosensory processing.." Behavioural brain research , vol. 207 , no. 2 , pp. 368-76.
M J Chacron, N Toporikova, Eric S. Fortune. 2009. "Differences in the time course of short-term depression across receptive fields are correlated with directional selectivity in electrosensory neurons.." Journal of neurophysiology , vol. 102 , no. 6 , pp. 3270-9.
Eric S. Fortune, M J Chacron. 2009. "From molecules to behavior: organismal-level regulation of ion channel trafficking.." PLoS biology , vol. 7 , no. 9 , pp. e1000211.
E M Hitschfeld, S A Stamper, K Vonderschen, Eric S. Fortune, M J Chacron. 2009. "Effects of restraint and immobilization on electrosensory behaviors of weakly electric fish.." ILAR journal , vol. 50 , no. 4 , pp. 361-72.
S Carver, E Roth, N J Cowan, Eric S. Fortune. 2008. "Synaptic plasticity can produce and enhance direction selectivity.." PLoS computational biology , vol. 4 , no. 2 , pp. e32.
N J Cowan, Eric S. Fortune. 2007. "The critical role of locomotion mechanics in decoding sensory systems.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 27 , no. 5 , pp. 1123-8.
J U Ramcharitar, E W Tan, Eric S. Fortune. 2006. "Global electrosensory oscillations enhance directional responses of midbrain neurons in eigenmannia.." Journal of neurophysiology , vol. 96 , no. 5 , pp. 2319-26.
Eric S. Fortune. 2006. "The decoding of electrosensory systems.." Current opinion in neurobiology , vol. 16 , no. 4 , pp. 474-80.
Eric S. Fortune, G J Rose, M Kawasaki. 2006. "Encoding and processing biologically relevant temporal information in electrosensory systems.." Journal of comparative physiology. A, Neuroethology, sensory, neural, and behavioral physiology , vol. 192 , no. 6 , pp. 625-35.
E W Tan, J M Nizar, E Carrera-G, Eric S. Fortune. 2005. "Electrosensory interference in naturally occurring aggregates of a species of weakly electric fish, Eigenmannia virescens.." Behavioural brain research , vol. 164 , no. 1 , pp. 83-92.
J U Ramcharitar, E W Tan, Eric S. Fortune. 2005. "Effects of global electrosensory signals on motion processing in the midbrain of Eigenmannia.." Journal of comparative physiology. A, Neuroethology, sensory, neural, and behavioral physiology , vol. 191 , no. 9 , pp. 865-72.
Eric S. Fortune, G J Rose. 2003. "Voltage-gated Na+ channels enhance the temporal filtering properties of electrosensory neurons in the torus.." Journal of neurophysiology , vol. 90 , no. 2 , pp. 924-9.
Eric S. Fortune, G J Rose. 2002. "Roles for short-term synaptic plasticity in behavior.." Journal of physiology, Paris , vol. 96 , no. 5-6 , pp. 539-45.
Eric S. Fortune, G J Rose. 2001. "Short-term synaptic plasticity as a temporal filter.." Trends in neurosciences , vol. 24 , no. 7 , pp. 381-5.
Eric S. Fortune, G J Rose. 2000. "Short-term synaptic plasticity contributes to the temporal filtering of electrosensory information.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 20 , no. 18 , pp. 7122-30.
G J Rose, Eric S. Fortune. 1999. "Frequency-dependent PSP depression contributes to low-pass temporal filtering in Eigenmannia.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 19 , no. 17 , pp. 7629-39.
G J Rose, Eric S. Fortune. 1999. "Mechanisms for generating temporal filters in the electrosensory system.." The Journal of experimental biology , vol. 202 , no. Pt 10 , pp. 1281-9.
Eric S. Fortune, G J Rose. 1997. "Passive and active membrane properties contribute to the temporal filtering properties of midbrain neurons in vivo.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 17 , no. 10 , pp. 3815-25.
Eric S. Fortune, G J Rose. 1997. "Temporal filtering properties of ampullary electrosensory neurons in the torus semicircularis of Eigenmannia: evolutionary and computational implications.." Brain, behavior and evolution , vol. 49 , no. 6 , pp. 312-23.
G J Rose, Eric S. Fortune. 1996. "New techniques for making whole-cell recordings from CNS neurons in vivo.." Neuroscience research , vol. 26 , no. 1 , pp. 89-94.
Eric S. Fortune, D Margoliash. 1995. "Parallel pathways and convergence onto HVc and adjacent neostriatum of adult zebra finches (Taeniopygia guttata).." The Journal of comparative neurology , vol. 360 , no. 3 , pp. 413-41.
D Margoliash, Eric S. Fortune, M L Sutter, A C Yu, B D Wren-Hardin, A Dave. 1994. "Distributed representation in the song system of oscines: evolutionary implications and functional consequences.." Brain, behavior and evolution , vol. 44 , no. 4-5 , pp. 247-64.
Eric S. Fortune, D Margoliash. 1992. "Cytoarchitectonic organization and morphology of cells of the field L complex in male zebra finches (Taenopygia guttata).." The Journal of comparative neurology , vol. 325 , no. 3 , pp. 388-404.
D Margoliash, Eric S. Fortune. 1992. "Temporal and harmonic combination-sensitive neurons in the zebra finch's HVc.." The Journal of neuroscience : the official journal of the Society for Neuroscience , vol. 12 , no. 11 , pp. 4309-26.
COLLAPSE
Other
"Electric fishes: neural systems, behaviour and evolution."
The Journal of experimental biology, July (3rd Quarter/Summer), 2013.
The Journal of experimental biology, July (3rd Quarter/Summer), 2013.